One goes to the transmission if it's automatic (vacuum diaphram on the A4LDs) one goes to the brake booster, one goes to the little plastic "pill" shaped thing on the passenger side engine compartment, One goes to the PCV, etc...I do have one capped off on mine.
Not all of the connections are used all of the time I have a few capped off. If you have a base model truck with a manual trans you could have quite a few that are blocked off and don't go anywhere.
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